A Statistical Analysis of the Social Behavior of the Male Stumptail Macaque (Macaca arctoides) Table I: Frequency of Behaviors as a Function of Dominance Status Page 20

Obtained frequency (a)
Behavior Abbrev. dominant
(vs.sub.)
dominant
(vs.dom.)
sub. Bertrand's terminology (b)
Behaviors shown mostly by dominant (vs. subordinates)
Hit, yank, shove or bump
Lying down
estrain firmly
Biting the back
Patrol locomotion
Restrain roughly
Run towards
Yawn
Round mouth frown
Walk away
HYSB
Lyng
Rstf
BtBk
Ptrl
RstR
RnTo
Fc4X
Fc21
WkAw
  19
  17
  10
  18
117
  24
  11
  37
  44
206
    0
    0
    0
    4 (6.9)
  22 (38)
  11 (19)
    5 (8.6)
  11 (19)
  16 (27)
  86 (147)
    0
    0
    0
    0
  23
    0
    1
  15
  24
125
Pat, slap, cuff, pull, grab, push
Same
Not distinguished
Not distinguished from BtFr
Patrolling
Not distinguished
See (c)
Same. See (d)
Not mentioned. See (d, e)
See (c)
Behaviors shown mostly by dominant (vs. dominant)
Teeth chomping
Biting-frontal
Masturbation
Threat lunge
Manipulate genitals
   of opponent
Autostimulation
Tremulo exhalation
Mounting
Lip smacking
Biting self
Teeth chattering face
Repeated bouncing
Bared-Teeth face
Grunting
Wide open mouth
Barking
Open mouth threat
Pant-grunting
Presenting
Chmp
BtFr
Mstb
Lnge
MnpO
 
Auto
Exhl
Mont
FcLS
BtSf
FcTC
RBnc
Fc23/33
Grnt
Fc3X
Bark
Fc31
Pant
Prst
   0
   2
 12
   1
   2

   7
 15
 30
 25
 20
 27
 35
 12
 55
 20
 18
   9
 11
 79
  13 (22)
  10 (17)
  73 (125)
    4 (6.9)
    5 (8.6)

  16 (27)
  31 (53)
  37 (63)
  73 (125)
  26 (45)
  52 (89)
  29 (50)
  47 (81)
  65 (111)
  56 (96)
  13 (22)
  22 (38)
  26 (46)
  65 (111)
    0
    0
  24
    1
    1

    3
  17
    9
  52
  10
  35
    1
  51
  35
  59
    4
  29
  23
  73
Not mentioned (f)
See biting the back
Same
Same
Genital manipulation

Same
Same
Same
Lipsmacking face
Not mentioned. See (g)
Same
Same
Bared teeth threat, fright faces
Several types of grunts
Not mentioned. See notes (d, e)
High pitched barks
Open mouth threat face
Same
Same
Behaviors shown mostly by subordinates
Crouching
Screaming
Biting cable
Hit at -no contact
Cooing
Run away
Climb up
Climb down
Self grooming
Grin or grimace
Allo-Grooming
Crch
Scrm
BtCb
HtNC
Coo
RnAw
ClbU
ClbD
SGrm
Fc13
AGrm
   0
   1
   3
   1
 12
   4
 17
 15
 92
   8
 66
    0
    2 (3.4)
    0
    0
    0
    7 (12)
    1 (1.7)
    1 (1.7)
  75 (129)
  13 (22)
  43 (74)
    9
  32
  22
    5
  25
  20
  33
  22
200
  24
  96
Same
Various types of screams
No restraint cable used
Intention pat, slap, cuff
Same
See (c)
Climbing
Climbing
Same
Grin face
Social grooming
Behaviors shown equally by animals of any status
Sitting
Walk towards
Standing
Walk parallel to other
Cage shaking
Sit
WkTo
Stnd
WkPp
Shkg
456
147
  55
  46
    5
205 (351)
  82 (141)
  63 (108)
  22 (38)
    0
516
185
  84
  62
    4
Various types of sitting
See (c)
Not distinguished
See (c)
Branch shaking

(a) Frequency that each behavior occurred by dominants versus subordinates, dominants versus dominants, and subordinates versus dominants. Frequencies of behaviors by dominants versus dominants are multiplied by 1.714 (in parentheses) to estimate the frequency that might have been expected if the total number of tests and observed behaviors had been equal in each category.

(b) Terminology follows that of Bertrand [1969] unless otherwise indicated.

(c) Terminology for running and walking is based only on speed and direction without regard for animal's apparent intention, unlike the system used by Bertrand.

(d) Static facial expressions were noted in terms of lip and mouth retraction (see text).

(e) Described in stumptall macaques by Chevalier-Skolnikoff [1974].

(f) Described in rhesus macaques as "teeth grinding."

(g) Described in stumptall macaques by Goosen [1975].

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